According to our current knowledge, the orangutan (Pongo pygmaeus) was the first great-ape species to split off from the evolutionary tree that led to modern humans. This happened about 14 million years ago (Goodman, 1999). In searching for the origins of human cognition and human language it is therefore imperative to gain knowledge about the cognitive and communicative abilities of this species. This includes the study of the intentional use of communicative signals like gestures [...]
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